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DiscussionThe results show that various aspects of the sitatungas’ behaviourare significantly different between biologically-relevant and otherenclosure zones. The frequencies of three behaviours (standing,sitting/ruminating and feeding) were significantly different frommorning to afternoon, thereby reflecting, in part, natural activitypatterns. The modified SPI value (0.61) indicates that the sitatungawere not utilising all aspects of their enclosure equally and the sitatungas’significant preference for reedy / long-grassy areas is supportedby the inferential analysis and by Figure 5. The herd spent54% of its time in only 28% of the enclosure, suggesting that improvementsto the remaining 72% encourage more widespreadmovement and a fuller development of a daily activity pattern.The increased usage of these natural areas may also be explainedby the behavioural ecology of the sitatunga and the need toconduct important state behaviours (feeding, resting, standing) in anenvironment most suitable for their performance. Comparing withobservations from Owen (1970), Kingdon (1982), Games (1983),Estes (1991), Starin (2000) and Robinchaud (2011), sitatunga wereobserved to remain sedentary in reedy, or long-grass beds duringthe day; such habitats were reflected in this captive setting and thesame behaviours were observed within them. Important functionsof long-grass for sitatunga in the wild include places to hide fromPredator, penyediaan tempat berlindung dari suhu tinggi dan situs untukperenungan (Estes 1991; Wronski et al., 2006). Gambar 4 menunjukkanrelatif proporsi waktu yang dihabiskan di daerah yang relatif kecilkandang (11%) dibandingkan dengan Semua Ruang lain yang tersedia untukhewan. Sementara tidak semua fungsi perilaku dibutuhkan per se dalamtawanan pengaturan, sangat penting untuk memberikan binatang dengan pilihan untukmelakukan perilaku yang memiliki motivasi tinggi nilai (Duncan1998); aktivitas tersebut dapat diandalkan indikator positif kesejahteraan(de Jonge et al., 2008). Dianjurkan sitatunga yang diberikandaerah-daerah yang lebih panjang rumput dan alang-alang untuk memfasilitasi preferensi mereka jelasuntuk jenis "habitat".
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