The prehistoric peoples of Wales left behind megaliths and other impressive monuments. They were followed by settlements of Celts in the region. The Romans occupied the region from the 1st to the 5th century A.D. Thereafter Angles, Saxons, and Jutes invaded the British island, but they left Wales virtually untouched. Beginning in the 8th century, the various Welsh tribes fought with their Anglo-Saxon neighbors to the east, but the Welsh were able to thwart attempted invasions. After William the Conqueror subdued England in 1066, however, his Norman armies marched into Wales in 1093 and occupied portions of it. By 1282, the English conquest of Wales was complete, and in 1284, the Statute of Rhuddlan formalized England's sovereignty over Wales. In 1301, King Edward I gave his son, who later became Edward II, the title Prince of Wales, a gesture meant to indicate the unity and relationship between the two lands. With the exception of Edward II, all subsequent British monarchs have given this title to their eldest son.