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The ingestion of seeds by earthworms alters their germinationthrough the physical damage of the seed coat in the earthwormgizzard or through their partial or total digestion in the earthwormgut (Grant, 1983; McRill and Sagar, 1973). The partial damage ofseeds might favor seed germination by breaking seed dormancy(Eisenhauer et al., 2009; Shumway and Koide, 1994). The impact ofearthworms on seeds might be driven by their preferences forcertain seed species over others This preference is mainlydriven by seed traits, which is the case for small and oil-rich seedsof Origanum vulgare and Urtica dioica (Aira and Piearce, 2009;Clause et al., 2011; Eisenhauer et al., 2009).
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